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After a few scuffles with these mutants, the turtles find that these mutants were turned into their current state by past experiments of Shredder's scientists.

They follow the tunnel and they are confronted by genetically mutated humans. Donatello's crystal start glowing and the turtles decide to investigate. Raphael kicks Michelangelo into a wall, revealing a large tunnel behind it. Afterward, while Raphael and Michelangelo spar for fun, Donatello analyzes a strange crystal he found in their home, noting that they look like mutated brain cells. Baxter Stockman soon uses invisible foot tech ninjas to capture Raphael, and Donatello is forced to rescue him. At the surface, he is confronted by Purple Dragon thugs, Casey Jones, and Dragonface. Michelangelo eventually gets on Raphael's nerves, making Raphael leave to the surface. Shortly after a group of mouser robots destroy the turtles' old home, they begin to look for a new home. The main gameplay loosely adapts the following season one episodes: 'Things Change', 'A Better Mouse Trap', 'Attack of the Mouser's, 'Meet Casey Jones', 'Nano', 'Darkness on the Edge of Town', 'The Way of Invisibility', 'Notes From the Underground' (Parts 1-3), and 'Return to New York' (Parts 1-3), as well as a level that is not derived from the animated series at all. A Challenge mode is unlockable by defeating Oroku Saki with any Turtle in the Story Mode, which needs to be complete to unlock Hamato Yoshi and his dojo.

In the versus mode, players can fight as all 4 turtles, Splinter, Casey Jones, Hamato Yoshi, The Turtlebot, Hun, Oroku Saki, and The Shredder. There is a story mode for one or two players, and there is also a versus mode where two players can fight head to head. Each turtle has his own unique set of levels to complete. The player can play as either Leonardo, Donatello, Michelangelo or Raphael.

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